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Crop Yields

Crop yields
Crop yield is the most widely used parameter in assessing the desirability of an alley cropping system (Nair 1993). There is increasing information on the effect of alley cropping with various hedgerow species on crop yields in various climatic and edaphic conditions. Results of studies on Alfisols and other high base status soils in the humid and subhumid tropics have usually been more promising in terms of crop yield. Soil nutrient status is better and tree-crop competition in the topsoil is less severe. For example, Kang et al. (1990) showed in an alley cropping study in Nigeria that maize yield, after eight years of rotation with cowpea, could be maintained at 2 ton ha-1 using Leucaena leucocephala prunings alone, improved to over 3 ton ha-1 using prunings supplemented by fertilizer (80 kgN ha-1), comparing to 0.66 ton ha-1 in the no-pruning, no-fertilizer control. An earlier study by Kang et al. (1981) showed that applying 10 ton ha-1 of fresh L. leucocephala prunings had the same effect on maize yield as applying 100 kg ha-1 of nitrogen fertilizers; however, such high amount of pruning material had to be supplemented by external sources. Some results are less successful. For example, in a study of alley cropping with G. sepium and L. leucocephala in Nigeria, Lal (1989a) found a large decline in the yields of maize (60% of the initial year) and cowpea (31%) after six years.

Trials in the humid zone on acid and low base status soils have given mixed, often less successful results. Nutrient availability is usually low in the soils, and root competition likely intensified due to acidity and toxicity of subsoils. For example, in an alley cropping study with Inga edulis on an infertile acid Ultisol in Peru, Fernandes et al., (1993a) found grain yields of rice under alley cropping (with and without fertilizer) declined considerably over five rice crops to less than 0.8 ton ha-1, compared to the steady increase and stabilization of rice yields of over 1.6 ton ha-1 in the fertilized control plot. The low yields under the alley cropping treatment were attributed to tree-crop root competition, where reduced crop yields due to root competition between hedgerows and crops were detected at 11 months after hedgerow establishment.

Similar low yields have been reported on a highly weathered Oxisol in Sumatra by Evensen et al. (1995), where yields of rice and cowpea alley-cropped with P. falcataria, C. calothyrsus and G. sepium declined over four years to unacceptable low levels (cowpea) or to crop failure (rice), until fertilizer inputs were increased and G. sepium replaced with F. macrophylla because of poor growth. Thereafter, crops yields increased and returned to original levels. Soil cations showed a similar pattern of change. Evensen et al. (1995) concluded that there was little build-up of nutrient cations due to recycling by trees, and that successful alley cropping on acid soils required maintenance of soil fertility with external inputs.

Selection and husbandry of hedgerows
Three aspects of alley cropping provide the most opportunity for management to improve efficiency and profitability of the system: tree species selection, hedgerow husbandry, interhedgerow spacing. As mentioned earlier, fast-growing leguminous perennials with deep and non-spreading rooting pattern are often preferred to attain higher pruning biomass, soil nitrogen status enhancement, nutrient uptake from subsoils, and reduced tree-crop root competition. Depending on the growth vigor and decomposition characteristics of prunings obtained from selected hedgerow species, the shoot pruning regime can be adjusted to minimize shading and achieve better synchrony between nutrient release from mulch/green manure and peak crop nutrient demand, and hence nutrient use efficiency. Decomposition rate and nutrient release pattern can also be adjusted by modifying the method of application of the prunings as mulch or green manure.

Nutrient release is not the only concern when selecting the decomposition characteristics of hedgerow species. In crop lands where weed infestation or soil erosion presents a particularly serious problem, providing a good ground cover for a longer period might be more important than the quick provision of nutrients. In such cases prunings with relatively rapid decomposition rates from species such as Leucaena leucocephala, Gliricidia sepium and Erythrina spp. might not be as desirable as slower-decomposing mulches of species such as Cassia siamea, F. macrophylla and Dactyladenia (syn. Acioa) barteri. MacLean (1992) suggested that diversified hedgerows with function-specific species may be more effective than hedgerows with single "multipurpose" tree species, especially in situations where erosion control and weed suppression are of primary concern.

Interhedgerow spacing
Several conflicting objectives need to be considered when determining the optimal interhedgerow spacing. Closer interhedgerow spacing, i.e. more hedgerows, can provide higher pruning biomass and nutrient yields per unit area in an alley cropping system. The nutrient cycling benefits of trees can also be more prominent. Weed suppression between cropping cycles is more effective with narrower alley where rapid canopy closure is possible. However, tree-crop competition for resources (solar radiation, moisture and nutrients) is intensified with smaller interhedgerow spacing. Higher proportion of tree-crop interface -- areas with the most pronounced depression on crop yield -- exists on crop land with more hedgerows. Planting double hedgerows can reduce interface while providing pruning biomass at a level similar to that of single hedgerows with smaller interhedgerow spacing. Based on the results of various on-station and on-farm trials, and on farmers' acceptance (mainly based on ease of manual cultivation), an interhedgerow spacing of 4 to 6 m was observed to be optimal for species such as G. sepium and L. leucocephala in the humid tropics (Kang 1993).

Where alley cropping is used for erosion control on sloping lands, there is less scope for adjustment of interhedgerow spacing. The introduction of trees on arable land also reduces the land area available for crop production. This reduction of land devoted to producing food crops may or may not translate into reduction of total crop yields, depending on the effectiveness of the system to improve yield per land cropped through nutrient provision and weed suppression. Studying yields of taro alley-cropped with Calliandra calothyrsus and G. sepium, in Western Samoa, Rosecrance et al. (1992) found that, even though 4-m interhedgerow spacing was most effective in suppressing weed and a slight yield increase over the control was found on a per plot basis in the alleys, total taro yields were significantly reduced in the 4-m alleys (6 ton ha-1) compared to the 5-m, 6-m, and control plots (8 ton ha-1). These yield reductions were primarily a result of more land taken out of taro production and put into hedgerow growth.

External inorganic fertilizer input
In alley cropping systems, the annual removal of nutrients is usually greater than in most agroforestry systems, resulting in greater demand on the system to supply and/or recycle nutrients. Nitrogen fixation by hedgerow leguminous trees accounts for the only significant nutrient addition from external source in an alley cropping system. Otherwise, prunings applied to alley crops are recycling, not augmenting, nutrients within the system and thus do no offset the nutrients lost via crop harvest. External inputs to the system from precipitation are small. Some nutrients, otherwise unavailable to crops because they are below the rooting zone of the crops, might be brought into the system from deeper layers in the soil by deep-rooting trees, but the magnitude of this "input" is not known (Nair 1993, Palm 1995).

Much of the research on alley cropping has addressed the ability of this system to maintain productivity with or without inorganic fertilization, in particular the capacity of leguminous species to substitute for nitrogen fertilizer application. A review of alley-cropped maize show that the application of inorganic nitrogen fertilizers considerably increases (up to 1000 kg ha-1) yields of maize alley cropped with L. leucocephala, but gives much smaller increases with G. sepium, F. macrophylla, or other alley cropping species (Szott and Kass, 1993). In another study on acidic soils, fertilizer additions (50 kg N, 25 kg P, 20 kg K, 35 kg Ca and 16 kg Mg ha-1) to rice and cowpeas alley cropped with Inga edulis resulted in significantly greater grain yields in the fertilized treatment in the fourth through the seventh crops of a seven-crop-long sequence (Fernandes 1990).

These results from fertilization studies indicate that nutrient availability can be a limiting factor in unfertilized alley crop systems. This is especially likely on acidic, infertile LAC soils in the humid tropics where the ability of hedgerow leguminous trees to recycle nutrients or fix nitrogen appears to be restricted by the intrinsically low levels of available nutrients and high levels of elements toxic to plant growth (Szott and Kass 1993). For example, nodulation by legumes can be prevented by deficiency of phosphorus, which results in low levels of nitrogen fixation and increased competition between trees and crops for available phosphorus. Balances of phosphorus additions and removals in alley cropping systems on infertile and acidic, or fertile but phosphorus- deficient soils suggest that phosphorus may eventually limit the productivity of these systems (Szott and Kass 1993), supporting the argument that pruning alone, especially on infertile soils, cannot sustain the productivity of continuous alley cropping because even if nitrogen demand can be met by biological nitrogen fixation, other nutrients, notably phosphorus, will ultimately become limiting after continuous exports (Szott 1991a, b, Evensen 1995).

Hedgerow perennials can be affected by the addition of inorganic fertilizers as well. For example, on an acidic soil in Rwanda, Yamoah et al. (1989) observed that growth of Sesbania sesban, Calliandra calothyrsus, Leucaena leucocephala and Markhamia lutea increased with manure additions of up to 5 Mg ha-1 in the first four months after transplanting, but by the eighth month there was no response to manure. Fernandes (1990) noted that pruning biomass and nutrient contents were greater in Inga edulis hedgerows bordering plots fertilized with N, P, K, Ca and Mg than those bordering unfertilized plots. Nutrient uptake from applied fertilizers by hedgerows can be seen as a benefit of more efficient nutrient cycling and retention, since less added nutrients will be leached and lost from the system; however, it also represents undesirable competition between trees and crops for nutrients intended for crop production. Nitrogen fertilization may also decrease the amount of nitrogen fixed by leguminous trees.

Introduction | Definition | Hedgerow Species I Species Selection Criteria | Soil Organic Matter & Nutrients | System Management | Crop Yields | Soil Conservation | Weed Dynamics | Tree-Crop Competition | References

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